$10,000,000+ (Contributor)
Bill & Melinda Gates Foundation [21]
Eli Lilly and Company [16] +

$5,000,000-$9,999,999 (Contributor)
Horizon Therapeutics plc [2] +
National Institutes of Health [14]
Novartis Pharmaceuticals Corporation [23]

$2,500,000-$4,999,999 (Contributor)
AbbVie Inc. [9]
Amgen Inc. [20]
Bayer AG [10]
Boehringer Ingelheim Pharmaceuticals, Inc. [5]
GlaxoSmithKline [24]
ImmunityBio [2]
Johnson & Johnson [22]
Merck Sharp & Dohme Corp. [25]
Pfizer Inc. [25]
Sanofi US Services Inc. [21]
Visterra
Wellcome [4]

$1,000,000-$2,499,999 (Contributor)
Biogen Inc. [12] ∞^
Bristol Myers Squibb [23]
Cytokinetics, Inc. [3]
Danaher Corporation
Genentech, A Member of the Roche Group [16]
Genmab US, Inc. [5]
Gilead Sciences, Inc. [7]
Ionis Pharmaceuticals, Inc. [3]
McKnight Brain Research Foundation [2]
Novo Nordisk A/S [5]
Regeneron Pharmaceuticals, Inc. [10]
Takeda Pharmaceutical Company Limited [15]
UCB, Inc. [4]
Ultromics

$500,000-$999,999 (Contributor)
Alzheimer's Association ® [18]
American Heart Association, Inc.
American Society of Echocardiography
AstraZeneca Pharmaceuticals LP [21]
Congress of Neurological Surgeons Foundation
Eisai Inc. [3]
Gates Ventures [4]
GENETHON
Otsuka Pharmaceutical Development & Commercialization, Inc. [3]
Ovid Therapeutics, Inc.
RTW Charitable Foundation
Syndax Pharmaceuticals Inc
Twist Bioscience Corporation +
Dr. and Mrs. Robert H. Wurtz [2]∆

$250,000-$499,999 (Contributor)
Estate of Michael T. Davis¥
Echosens SA [2]
GE Healthcare [2]
Cathy Graham [2]
Intercept Pharmaceuticals, Inc.
LGC Clinical Diagnostics
Spark Therapeutics [2]
Thermo Fisher Scientific Inc. [3]
Ultragenyx Pharmaceutical [3]

$100,000-$249,999 (Contributor)
Alkermes, Inc. [4]
Alzheimer's Drug Discovery Foundation [10]
American Psychiatric Association Foundation [3]
Amgen Foundation [2]
Arthritis Foundation [13]
Deeda Blair [7] ∞
BrightFocus Foundation [4]
Buffy and William Cafritz Family Foundation
CureDuchenne [2]
Foundation Fighting Blindness
IBM Corporation
Jazz Pharmaceuticals [3]
The Robert Packard Center for ALS Research at Johns Hopkins
Lupus Research Alliance [2]
Ann Lurie [11]
National Alliance on Mental Illness [4]
National Psoriasis Foundation [2]
NuProbe
One Mind [3]
The Benjamin Harry Peikin Foundation, Inc.
Pharmaceutical Research and Manufacturers of America [18]
REGENXBIO Inc. [2]
ROCHE
The Edmond J. Safra Foundation [5]
Sage Therapeutics [4]
Nina K. Solarz [3]
Somaiya Family Foundation
Sysmex [2]
Schizophrenia & Psychosis Action Alliance [2]
The Uhrig/Vournas Fund of Vanguard Charitable

$50,000-$99,000 (Contributor)
The ALS Association [2]
American Association for Dental Research [10]
The Association for Frontotemporal Degeneration [6]
Astellas Pharma Inc. [3]
Autoimmune Association
Day One Biopharmaceuticals
Dr. and Mrs. Marijn Dekkers [4]
Deloitte
Julie Louise Gerberding, M.D., M.P.H. and David Rose
Peter and Judy Kovler ∞
Mr. and Mrs. Paul M. Montrone [20] ∞
Novalis LifeSciences LLC [2]
Gilbert S. Omenn, M.D., Ph.D. and Martha A. Darling [12]
Ms. Virginia Schirrmeister and Mrs. Anne Goldrach
Fred A. and Donna Seigel [6]
Estate of James T. Wendel [2]¥

$25,000-$49,999 (Contributor)
Adaptive Biotechnologies Corporation [7]
Alector, Inc. [6]
Alliance for Regenerative Medicine [2]
American Society of Gene & Cell Therapy [2]
AMS Foundation for the Arts, Sciences and Humanities ∞
ASC Therapeutics, Inc.
Colgate-Palmolive Company [5]
Foundation Medicine, Inc.
Ikena Oncology [3]
Invicro [4]
Laboratory Corporation of America [3]
Sherry and Alan Leventhal Family Foundation
Mr. and Mrs. Joel S. Marcus ∞
Michael Jefferson Meagher [3]
National Rongxiang Xu Foundation
Annette L. Nazareth and Roger W. Ferguson, Jr. [4]
Oncoheroes Biosciences
Steven and Jann Paul [3] ∞
Rett Syndrome Research Trust [2]
Dame Jillian Sackler [20] ∞
Seagen, Inc. [2]
Skingenix, Inc.
The Grass Foundation
Drew Weissman, M.D.
Trust of Lillian S. White∆¥

$10,000-$24,999 (Contributor)
Anonymous (1) ∞^
The Jeffrey A. Abrams and Rosalyn L. Abrams Charitable Trust
ADx NeuroSciences [4]
American Endowment Foundation [5] ∞
American Society for Biochemistry and Molecular Biology
AMRA Medical [5]
ARUP Laboratories +
Berger Family Foundation, Inc.
Biognosys
Edward M. Blair, Jr.
Francis I. Blair
The Bluefield Project to Cure Frontotemporal Dementia [4]
Donald Burrows [3]
C₂N Diagnostics [5]
Clario. [15]
Clinical Ink
Eileen and Jack Connors, Jr.
Larry Corey, M.D.
Caroline M. Devine
James H. Donovan [11] ∞
Beatrice Forbes Herndon
Frederick National Laboratory for Cancer Research [6]
Friends of Cancer Research [13]
Fujirebio
GHR Foundation
Goulston & Storrs PC [4]
Estate of Jack Gramlich¥
Carol-Ann Harris [10]
Dr. and Mrs. Thomas R. Insel [6] ∞
InterVenn Biosciences
Arian Karbasi and Annie Coe ∞
Patricia S. Kohlen [4]¥
Julie Bell Lindsay [8]
Lumos Labs, Inc.
H. Lundbeck A/S [7]
Steve and Sherry Mayer [10] ∞
Mintz, Levin, Cohn, Ferris, Glovsky and Popeo, P.C.
MITRE Corporation
Morgan Stanley & Co. Incorporated [4]
The Multiple Myeloma Research Foundation [3]
Dr. and Dr. Gary J. Nabel
Bob and Sally Newcomb
NEWMARK [2]
Nordic Bioscience A/S [4]
Olink Proteomics [6]
Carl Page ∞
The Pew Charitable Trusts [9]
Posey-Glickert Foundation [5]
Prolaio, Inc.
Rhino Health
Saladax Biomedical, Inc.
Sapient Bioanalytics
Jane M. Sayer, Ph.D. [2]
Sengenics Corporation [4]
Gerald R. Sigal and Ellen V. Sigal, Ph.D. [3]
Sjögren's Foundation [4]
Solomon H. Snyder, M.D. [7] ∞
SomaLogic, Inc.
Russell W. Steenberg and Patricia Colbert [10]
Paul Stoffels, M.D. and Katelijne Bruurs [5] ∞
Trust of Anne Moore Strenk
Verily [2]
Vir Biotechnology
Windex Non-Toxic Settlement Fund ∞
The Richard H. Yearick Foundation [3] ∞
Elias A. Zerhouni, M.D. and Nadia Zerhouni, M.D. [8] ∞

$5,000-$9,999 (Contributor)
Anonymous (2)
Advanced Medical Technology Association - AdvaMed
Association for Molecular Pathology [5]
John Bertschy [4]
Ralph H. and Karen K. Craft [6] ∞
Jayne Koskinas Ted Giovanis Foundation for Health and Policy [6] ∞
Global Impact Combined Federal Campaign [19] ∞^
IMB Partners
Matthew Scher and Barbara Lazio [10] ∞
Lupus Foundation of America [2]
National Scleroderma Foundation
PathAI, Inc. [3]
Radiological Society of North America [14]
John and Katrina Rogers
Steven L. and Karin Siegel [2] ∞
Truist [12]
Walter Stephan Kovshik Sr. Memorial Fund for Lymphoma Research
Steve and Chris Wilsey [14]

$2,500 - $4,999 (Contributor)
Ronald and Barbara Berke [13]
Competition Dynamics ∞
Drs. Maria & Ernesto Freire [11] ∞
Randy K. Glantz [5]
Henry L. Hecht [6]
Dr. and Mrs. Paul L. Herrling [9] ∞
Host Hotels & Resorts, Inc. ∞^
Lisa Kuzel [3] ∞
Edison T. Liu, M.D., Ph.D. and Margaret B. Liu [5] ∞
Kunal Patel [3] ∞
Patricia Solvik Family Fund ∞
Lenore R. Salzman [7]
Jonathan G. Seidman, Ph.D. Christine E. Seidman, M.D. ∞
Society for Immunotherapy of Cancer [6]
Andrew and Elyse Steinhaus [7]
Sara Lou Whildin [10] ∞
Wolk Family Fund [5] ∞
Richard G. Wyatt, M.D. and Linda S. Wyatt, Ph.D. [8] ∞

$1,000 - $2,499 (Contributor)
Anonymous (4) ∞
ACT for NIH
The Air Products Foundation ∞^
American Association for the Advancement of Science
America's Charities [7] ∞^
Jeffrey D. and Ann Anderson [6] ∞
Dr. John P. and Andrea J. Atkinson
Baker-Merine Family Foundation ∞
Dr. Nadarajah Balasubramanian [6] ∞
Robert Balthaser and Ricardo C. Araneda, Ph.D. [6]
Joan and Brian Barrett
BlackRock ∞^
Margaret Blair [2]
Zachary T. Bloomgarden, Ph.D. and Kathy F. Bloomgarden, Ph.D. [2] ∞
Paula L. and William C. Bradley [4]
Marc and Debbie Breslawsky [3] ∞
Christopher Calingo ∞^
Dan Balliet and Jan Carlson [14] ∞
Gina D. Chalmers [4] ∞
Jeffrey Chow [5]
Stewart A. Daniels, M.D. [2] ∞
David Decker and Lisa Greenlees
Ronald Early [4] ∞
Elysium Health LLC
Alicia Emerson [6]
Exact Sciences
Faegre Drinker Biddle & Reath LLP
Joseph M. Feczko, M.D. and Leighton K. Gleicher
Jeffrey and Marilyn Finn [7] ∞
Friedman Family Gift Fund ∞
Keith Gendler [7] ∞
Matthew Giles
Leonard M. and Cynthia A. Glassman [5] ∞
Drs. David Golan and Laura Green ∞
E. Laura Goldberg and Michael D. Hamilton
Goodwin Procter, LLP
Google, Inc. [8] ∞^
Hewlett Packard Enterprise [5]
Eric Hirschhorn and Leah Wortham [12]
Susan C. Horowitz [8] ∞
Youngmi Ji, Ph.D. ∞
James W. Jones
Douglas M. and Lynn C. Klock [5] ∞
Teresa and Kevin A. Klock [8] ∞
Ronald L. Krall, M.D. and Susan J. Krall ∞
Debra R. Lappin, J.D.
John Madden, Jr. [4]
Dr. John and Stacy Martin [3]
Charles F. McCormick [2] ∞
Metcor/LSI
Raymond Michael [7] ∞
Christine Mina ∞^
National Cancer Registrars Association ∞
NextEra Energy, Inc.
Paul and Cindy Obermeyer [3] ∞
Mariann O'Brien [2] ∞
Susan Oldfield ∞
Sarah Palamara [5] ∞
Joseph G. Perpich, M.D., J.D. and Cathy J. Sulzberger ∞
Jeffrey Peterson [2]
Mr. and Mrs. Stephen Peth [14] ∞
Eric F. Polhamus [7] ∞
Sunny Raspet [10]
Gregory and Sherry Roper [1]
Sherry Roper ∞
Robert A. and Marjorie Rosenberg [10] ∞
John Salvino [2] ∞
Charles A. Sanders, M.D. and Ann E. Sanders [26] ∞
Barbara Santos [3]
Dr. John N. Sheagren [2] ∞
Robert B. and Tammy Renee Sher [3] ∞
Sheldon and Melissa Cohen Silberman
Richard I. and Anastasia Smith [10] ∞
Technomics Research [2] ∞
Christopher A. and Elizabeth Thoma [7] ∞^
William Morley and Caroline Trahan [6]
Jon and Kristin Vaver [15] ∞
Susan M. Weinert
Katie and Ted Wells [7] ∞
David Wholley [11] ∞
Julie and Howard Wolf-Rodda [16] ∞

$500-$999 (Contributor)
Anonymous (2) ∞
Abbott [2] ∞^
Jayne Abshire ∞
AcademyHealth
Dr. James S. Alexander [2] ∞
Stephen and Sharon Alpert [6] ∞
Amazon Smile Foundation [9] ∞
Apple Inc. [6] ∞^
Dennis A. Ausiello, M.D. ∞
Sheila Bassett [3] ∞^
Jon H. Beusen and Denise D. Beusen, Ph.D. [9] ∞
James K. and Deborah M. Bieging [6] ∞
Wolf and Lynn Blitzer ∞
Alex and Drew Burrows [3]
Erin Claywell
John F. Malmros and Rosemary F. Contin [3] ∞
Daniel Cunningham ∞
Darby Fund ∞
B.A. Dass [4]
Paul J. Davis, M.D. [3] ∞
F5 Networks [4] ∞^
Facebook Donors [6] ∞
Faye Fager [9] ∞
James M. Felser, M.D. [13]
Joseph N. and Michie Flanz [7]
We Chen Foo [3] ∞
Stephen J. and Joyce A. Ford ∞
Laren Friedman [9] ∞
Gavelli Generational Gift Fund
Randall and Holly Griffin [6] ∞
Robert M. and Susie Hathaway
Chris and Laura Hazzard [14]
Mary Hennessey and Daniel Cunningham ∞
Ryan M. Hohman
John L. and Mary Emma C. Hoye [8]
Hubble Charitable Fund [3] ∞
Luanne Hudson [2] ∞
International Monetary Fund [3] ∞^
Terrence J. Jach, Ph.D. ∞
James F. and Gudrun Jeffrey [7] ∞
Richard Jonas and Katherine Vernot-Jonas [15] ∞
Michael M. Kaback, M.D. [2] ∞
Charles E. Kaufman Medical Fund [7]
Jane Krmpotich ∞
Frances Krogstad ∞
Lafayette 89 [4]
John Larabee [5] ∞
Aleah Laxton [5] ∞^
Janelle Lewis [3]
Catherine Master [6] ∞
Dr. Donald R. Mattison ∞
The Honorable and Mrs. Matthew McHugh [7]
Microsoft Corporation [7] ∞^
Jonathan R. Oates [5] ∞
Robyn and Matt Painter [4]
Farhan Panthaki [5] ∞
Brian Park [5] ∞
Marilyn H. Paul
Anne Pfaelzer De Ortiz ∞
Susan Butler Plum
The Relias Family [8]
Research!America
Barbara J. Richmon ∞
Julia Robbins ∞
Jennifer Rosenbluth-Stoll and Peter Stoll [5]
Darren Schneider [8] ∞
Ben Schwartz and Katie Kennedy [3] ∞
Sealy & Company ∞
Daniel R. Shively [2] ∞
Jennifer Seigel ∞
Matthew M. Stayman
Lorene Steinberg [6]
Dr. Joan A. Steitz ∞
Bhuvaneswari Subramanian
Suncor Energy [3] ∞^
The Pittsburgh Foundation [4]
Dr. Samuel O. Thier [17]
Tischfield Family Charitable Gift Fund [2] ∞
William, Zani and Aycen Tolentino [14]
Steve and Robin Toomey
Paula J. Warrick, Ph.D. [4] ∞
Mark Youngblood and Todd Youngblood
Lucas and Katrina Yun-Nikolac [9] ∞^
Dr. Stuart H. Yuspa and Eleanor H. Yuspa [2]

$250-$499 (Contributor)
Jonathan Abney [3]
Joan Beck [12]
Albert M. Bernath, Jr., M.D. [2]
Will and Berta Blades [7]
Jonathan and Heidi Blythe [3]
William and Nancy Bonner
Walt and Jane Brandt [6]
Jacqueline G. Breen
Manson K. Brown [4]
Louis Maximilian Buja, M.D. [4]
Keith F. and Alison Burrows [4]
Joonhao Chuah [3]
James Collison
Deborah Cooper
Khue Dang
Diane Daum
Larry Day [7]^
Marla Diamond
Bob and Debra Ekman
Renee D. Enright
Margaret Faust
Arlene L. Feit [10]
Fitch Group [2]^
Shannon and David Forsell [2]
J. Michael Foster
Jason and Gloria Garver^
Thomas Gasparini [6]
Martin Gellert, Ph.D. [6]
Daniel and Bernice Goldberg
Christina Graham
Spencer Griffin
Stacy D. Harrison
Eric J. and Susan Hatch [6]
Kelley Hearne
Thomas Heefner [2]
Stephen and Teri Herbst [5]
Todd Hillman [3]
David B. and Janet S. Hoffberger
Stephanie L. James, Ph.D. [12]
Robert S. and Raissa H. Johnson [4]
Kye Kim
Dr. Richard Krauzlis [2]
Paul and Nancy Kurland
Melissa Kuskin [3]
Hale G. Lamont-Havers [4]
Philip and Nancy N. Lee [3]
Dr. and Mrs. Lewis A. Lipsitz [6]
Thomas A. and Nancy I. Lusk [6]
William and Stephanie Marra [4]
Anne Alexander Marshall, Ph.D. and Davis Marshall [18]
James J. and Marilyn P. McCullough
Randall McEntire [2]
Laurence McMillan [4]
Cheryl L. Melencio [4]
Veronika Memetova [2]
Judith Miller [5]
Jake Morgan
Mr. and Mrs. WW Dabney Charitable Fund
Gretchen H. Naylor [2]
Northrop Grumman Corporation [4]
NVIDIA Corporation^
Marshall and Kathy Peterson [7]
Robert and Paula Petterson [8]
PointOne Holdings Management
Jan and William Price [2]
Dr. Thomas C. Quinn
Kathleen M. Reardon [2]
Peny Resnick
Glen Richards [5]
Lori A. Rolnick [5]
Sidney Rosenzweig [9]
Dr. and Mrs. George Schneider [13]
Dorina Sepulveda [6]
Wonsock Shin, M.D.
Robert B. Sollitto
Drs. Byoung J. and Minkyung Song
Minkyung Song, Ph.D.
Michael Sullivan [7]
Anthony Tassone [10]
The Sunday Show Group
Lois and Dilawar Uthman
Susan Walters
Thomas Wibe
Ingrid Wiley [14]
Jay Yarington
Ying Cheung Yee
Joel M. Yesley [15]